A method of operating a nuclear fission reactor, the reactor comprising a reactor core, and a coolant tank containing coolant, the reactor core comprising an array of fuel assemblies arranged in generally parallel rows, each fuel assembly comprising one or more fuel tubes containing fissile fuel. For each row of the array, one or more spent fuel assemblies are removed from the array at a second end of the row, fuel assemblies are moved along the row from a first end to the second end; and one or more fuel assemblies are introduced to the array at the first end of the row. Each fuel assembly remains within a single row while the fuel assembly is within the array. At least the fuel-filled portions of the fuel tubes of each fuel assembly are immersed in the coolant while the fuel assembly is within the array.